@import url("../shared_base.css");

html {
  font-size: 100%;
  scroll-behavior: smooth;
}

:root {
  --bg: #eef5f0;
  --panel: #ffffff;
  --panel-soft: #f6fbf8;
  --text: #13251d;
  --muted: #5c6f67;
  --accent: #1f7a5d;
  --accent-strong: #0d4f3f;
  --accent-soft: #dff1e8;
  --line: #d5e4dd;
  --signal: #7fa8ff;
  --shadow: 0 18px 48px rgba(19, 37, 29, 0.1);
  --container-max: var(--container-lg, 82rem);
  --radius: 0.8rem;
  --radius-sm: 0.55rem;
  --radius-pill: 999rem;
}

*,
*::before,
*::after {
  box-sizing: border-box;
  margin: 0;
  padding: 0;
}

body {
  background:
    radial-gradient(
      circle at top left,
      rgba(31, 122, 93, 0.12),
      transparent 26%
    ),
    radial-gradient(
      circle at bottom right,
      rgba(127, 168, 255, 0.1),
      transparent 24%
    ),
    linear-gradient(180deg, #f4f8f5 0%, #ffffff 100%);
  color: var(--text);
  font-family: Arial, Helvetica, sans-serif;
  font-size: 1rem;
  line-height: 1.6;
}

a {
  color: inherit;
}

.page-hero {
  position: relative;
  min-height: clamp(24rem, 58svh, 36rem);
  overflow: hidden;
  background: #183b2c;
  color: #f3fff8;
  isolation: isolate;
}

.hero-backdrop {
  position: absolute;
  inset: 0;
  width: 100%;
  height: 100%;
  object-fit: cover;
  filter: saturate(1.01) contrast(1.03) brightness(1.02);
  transform: scale(1.01);
  z-index: var(--z-base, 0);
}

.hero-overlay {
  position: absolute;
  inset: 0;
  z-index: var(--z-overlay, 1);
  pointer-events: none;
  background:
    linear-gradient(
      90deg,
      rgba(3, 16, 13, 0.52) 0%,
      rgba(3, 16, 13, 0.3) 48%,
      rgba(3, 16, 13, 0.52) 100%
    ),
    linear-gradient(180deg, rgba(3, 16, 13, 0.12) 0%, rgba(3, 16, 13, 0.4) 100%);
}

.hero-shell {
  position: relative;
  z-index: var(--z-content, 2);
  width: min(var(--container-max), calc(100% - var(--container-gutter-tight, clamp(1rem, 4vw, 2rem))));
  min-height: inherit;
  margin: 0 auto;
  padding: clamp(0.85rem, 2.2vw, 1.35rem) 0;
  display: grid;
  grid-template-columns: minmax(0, 1.25fr) minmax(17rem, 0.75fr);
  gap: clamp(1rem, 3vw, 2rem);
  align-content: center;
  align-items: start;
}

.hero-copy {
  max-width: 64ch;
}

.hero-copy-body {
  display: grid;
  gap: 1rem;
}

.hero-copy-body > div {
  padding: clamp(0.6rem, 1.2vw, 0.95rem);
  border: 1px solid rgba(215, 228, 215, 0.2);
  border-radius: var(--radius-sm);
  background: linear-gradient(
    180deg,
    rgba(3, 10, 5, 0.5) 0%,
    rgba(3, 10, 5, 0.34) 100%
  );
  backdrop-filter: blur(6px);
}

.eyebrow,
.section-kicker {
  margin: 0 0 0.75rem;
  color: #9de9cd;
  font-size: 0.8rem;
  font-weight: 700;
  letter-spacing: 0.12em;
  text-transform: uppercase;
  text-align: left;
  white-space: normal;
}

.section-kicker {
  color: var(--accent);
}

h1,
h2,
h3,
p,
li,
dt,
dd,
th,
td {
  letter-spacing: 0;
}

h1 {
  margin: 0;
  font-size: clamp(1rem, 6vw, 5.2rem);
  line-height: 0.98;
  text-wrap: balance;
}

.hero-lead {
  margin: 1rem 0 0;
  color: rgba(243, 255, 248, 0.84);
  font-size: clamp(1rem, 0.7vw + 0.92rem, 1.25rem);
  line-height: 1.6;
  hyphens: none;
}

.subtitle {
  margin: 1rem 0 0;
  color: rgba(243, 255, 248, 0.84);
  font-size: clamp(1rem, 0.72vw + 0.9rem, 1.2rem);
  line-height: 1.6;
  hyphens: none;
  text-wrap: pretty;
}

.thesis-banner {
  margin-top: 1.25rem;
  padding: 1rem 1rem 1.05rem;
  border: 1px solid rgba(157, 233, 205, 0.18);
  border-radius: var(--radius);
  background: linear-gradient(
    180deg,
    rgba(6, 31, 23, 0.84),
    rgba(3, 18, 15, 0.92)
  );
  box-shadow: var(--shadow);
  backdrop-filter: blur(10px);
}

.thesis-banner h2 {
  margin: 0 0 0.45rem;
  color: #f3fff8;
  font-size: clamp(1.15rem, 1vw + 1rem, 1.45rem);
}

.thesis-banner p {
  margin: 0;
  color: rgba(243, 255, 248, 0.82);
  line-height: 1.55;
}

.section-nav {
  display: flex;
  flex-wrap: wrap;
  gap: 0.625rem;
  margin-top: 1.25rem;
}

.section-nav a {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 2.75rem;
  padding: 0.5rem 1rem;
  border: 1px solid rgba(243, 255, 248, 0.3);
  border-radius: var(--radius-pill);
  background: rgba(4, 18, 15, 0.66);
  color: #f3fff8;
  text-decoration: none;
  font-size: 0.92rem;
  backdrop-filter: blur(8px);
  hyphens: none;
}

.section-nav a:hover,
.section-nav a:focus-visible {
  border-color: rgba(157, 233, 205, 0.55);
  background: rgba(13, 79, 63, 0.62);
}

.hero-panel {
  padding: 1rem;
  border: 1px solid rgba(243, 255, 248, 0.18);
  border-radius: var(--radius);
  background: rgba(4, 18, 15, 0.72);
  box-shadow: var(--shadow);
  backdrop-filter: blur(10px);
}

.summary-grid {
  display: grid;
  gap: 0.85rem;
}

.summary-block {
  padding-bottom: 0.85rem;
  border-bottom: 1px solid rgba(243, 255, 248, 0.14);
}

.summary-block:last-child {
  padding-bottom: 0;
  border-bottom: 0;
}

.summary-block h3,
.summary-block .summary-label {
  margin-bottom: 0.35rem;
  color: #9de9cd;
  font-weight: 700;
}

.summary-block p {
  margin: 0;
  color: rgba(243, 255, 248, 0.88);
  line-height: 1.45;
  hyphens: none;
}

.hero-panel dl {
  display: grid;
  gap: 0.85rem;
  margin: 0;
}

.hero-panel div {
  padding-bottom: 0.75rem;
  border-bottom: 1px solid rgba(243, 255, 248, 0.14);
}

.hero-panel div:last-child {
  padding-bottom: 0;
  border-bottom: 0;
}

.hero-panel dt {
  color: #9de9cd;
  font-size: 0.78rem;
  font-weight: 700;
  text-transform: uppercase;
}

.hero-panel dd {
  margin: 0.2rem 0 0;
  color: rgba(243, 255, 248, 0.88);
  line-height: 1.45;
}

main,
footer {
  width: min(var(--container-max), calc(100% - var(--container-gutter-tight, clamp(1rem, 4vw, 2rem))));
  margin: 0 auto;
}

main {
  padding: 0 0 3rem;
}

section {
  padding: 2rem 0;
  border-top: 1px solid var(--line);
}

section:first-of-type {
  border-top: 0;
}

h2 {
  margin: 0 0 0.85rem;
  color: var(--accent-strong);
  font-size: clamp(1.45rem, 1vw + 1.15rem, 2rem);
  line-height: 1.16;
}

h3 {
  margin: 0 0 0.6rem;
  color: var(--text);
  font-size: 1.05rem;
}

p,
li,
td,
th {
  font-size: clamp(1.02rem, 0.18vw + 0.98rem, 1.12rem);
  line-height: 1.65;
  text-align: justify;
  text-wrap: pretty;
  hyphens: auto;
}

p {
  margin: 0 0 1rem;
}

:where(main, .hero-panel) * {
  text-align: justify;
}

:where(main, .hero-panel)
  :where(
    h1,
    h2,
    h3,
    th,
    td,
    caption,
    .section-nav a,
    .action-link,
    .evidence-kicker,
    .evidence-tag,
    .metric-value,
    .metric-label,
    button
  ) {
  text-align: left;
  hyphens: manual;
}

code {
  padding: 0.1rem 0.28rem;
  border-radius: 0.35rem;
  background: rgba(31, 122, 93, 0.1);
  color: var(--accent-strong);
  font-size: 0.94em;
}

.card-grid {
  display: grid;
  grid-template-columns: repeat(auto-fit, minmax(14rem, 1fr));
  gap: 0.875rem;
  margin-top: 1rem;
}

.info-card,
.note-panel,
.figure-panel {
  padding: 1rem;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel);
  box-shadow: 0 10px 26px rgba(19, 37, 29, 0.06);
}

.info-card p {
  margin-bottom: 0;
  color: var(--muted);
}

.section-split {
  display: grid;
  grid-template-columns: minmax(0, 1.08fr) minmax(18rem, 0.92fr);
  gap: 1rem;
  align-items: start;
}

.classification-layout {
  display: grid;
  grid-template-columns: minmax(0, 0.9fr) minmax(0, 1.1fr);
  gap: 1rem;
  margin-top: 1rem;
  align-items: stretch;
}

.classification-tree {
  padding: 1rem;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: linear-gradient(180deg, #ffffff 0%, #f7fbf8 100%);
  box-shadow: 0 10px 24px rgba(19, 37, 29, 0.05);
}

.classification-tree p {
  margin: 0;
  color: var(--accent-strong);
  font-family: ui-monospace, SFMono-Regular, Menlo, Consolas, monospace;
  text-align: left;
  hyphens: none;
}

.classification-tree p + p {
  margin-top: 0.45rem;
}

.evolutionary-tree {
  margin: 1rem 0;
  border-left: 0.3rem solid var(--accent);
  overflow: hidden;
}

.evolutionary-tree .tree-root {
  margin: 0 0 0.9rem;
  color: var(--accent-strong);
  font-family: ui-monospace, SFMono-Regular, Menlo, Consolas, monospace;
  font-size: 1rem;
  line-height: 1.35;
}

.lineage-tree,
.lineage-tree ul {
  display: grid;
  gap: 0.7rem;
  min-width: 0;
  margin: 0;
  padding: 0 0 0 1.15rem;
  border-left: 1px solid var(--line);
  list-style: none;
}

.lineage-tree ul {
  margin-top: 0.7rem;
}

.lineage-tree li {
  position: relative;
  min-width: 0;
  font-size: 1rem;
  line-height: 1.45;
  text-align: left;
  hyphens: none;
}

.lineage-tree li::before {
  position: absolute;
  top: 1.05rem;
  left: -1.15rem;
  width: 0.85rem;
  border-top: 1px solid var(--line);
  content: "";
}

.lineage-node {
  display: grid;
  grid-template-columns: minmax(8.5rem, max-content) minmax(0, 1fr);
  gap: 0.2rem 0.8rem;
  min-width: 0;
  padding: 0.55rem 0.7rem;
  border: 1px solid rgba(213, 228, 221, 0.8);
  border-radius: var(--radius-sm);
  background: rgba(255, 255, 255, 0.72);
}

.lineage-node strong,
.lineage-node code,
.lineage-node span {
  min-width: 0;
  color: var(--accent-strong);
  overflow-wrap: break-word;
  word-break: normal;
  text-align: left;
  text-wrap: pretty;
  hyphens: none;
}

.lineage-node code {
  align-self: start;
  justify-self: start;
}

.source-metadata {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 0.75rem;
  margin: 1rem 0;
}

.source-metadata div {
  padding: 0.75rem;
  border: 1px solid var(--line);
  border-radius: var(--radius-sm);
  background: var(--panel-soft);
}

.source-metadata dt {
  color: var(--accent-strong);
  font-size: 0.78rem;
  font-weight: 700;
  letter-spacing: 0.06em;
  text-transform: uppercase;
}

.source-metadata dd {
  margin-top: 0.2rem;
  color: var(--muted);
}

.reactome-record .source-line a {
  color: var(--accent-strong);
  font-weight: 700;
}

.callout {
  margin: 1rem 0;
  padding: 1rem;
  border: 1px solid rgba(31, 122, 93, 0.16);
  border-radius: var(--radius);
  background: linear-gradient(180deg, #ffffff 0%, #f7fbf8 100%);
  box-shadow: 0 10px 24px rgba(19, 37, 29, 0.05);
}

.callout h3 {
  color: var(--accent-strong);
}

.callout p:last-child {
  margin-bottom: 0;
}

.tag-row {
  display: flex;
  flex-wrap: wrap;
  gap: 0.45rem;
  margin: 0 0 0.8rem;
}

.tag {
  display: inline-flex;
  align-items: center;
  min-height: 1.9rem;
  padding: 0 0.65rem;
  border-radius: var(--radius-pill);
  border: 1px solid var(--line);
  background: var(--panel-soft);
  color: var(--accent-strong);
  font-size: 0.8rem;
  font-weight: 700;
  text-transform: uppercase;
  letter-spacing: 0.06em;
}

.data-table {
  width: 100%;
  border-collapse: collapse;
  margin-top: 1rem;
  background: var(--panel);
}

.table-scroll {
  overflow-x: auto;
  margin-top: 1rem;
  border-radius: var(--radius);
}

.table-scroll:focus-visible {
  outline: 3px solid rgba(31, 122, 93, 0.35);
  outline-offset: 0.2rem;
}

.external-record {
  margin-top: 1rem;
  padding: 0.75rem;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel);
  box-shadow: 0 10px 26px rgba(19, 37, 29, 0.06);
}

.external-record-summary {
  display: grid;
  grid-template-columns: minmax(0, 1fr) auto;
  gap: 1rem;
  align-items: center;
  padding: clamp(0.9rem, 2vw, 1.25rem);
  border: 1px solid var(--line);
  border-radius: var(--radius-sm);
  background: var(--panel-soft);
}

.external-record-summary h3 {
  margin-bottom: 0.35rem;
  color: var(--accent-strong);
}

.external-record-summary p:not(.section-kicker) {
  max-width: 54rem;
  margin: 0;
  color: var(--muted);
}

.external-record-actions {
  display: flex;
  flex-wrap: wrap;
  gap: 0.55rem;
  justify-content: flex-end;
}

.record-link {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 2.75rem;
  padding: 0.5rem 0.85rem;
  border: 1px solid var(--accent-strong);
  border-radius: var(--radius-pill);
  color: var(--accent-strong);
  font-weight: 700;
  line-height: 1.2;
  text-align: center;
  text-decoration: none;
}

.record-link-primary {
  background: var(--accent-strong);
  color: #ffffff;
}

.record-link:focus-visible {
  outline: 3px solid rgba(31, 122, 93, 0.35);
  outline-offset: 0.2rem;
}

.external-record-note {
  margin: 0.75rem 0 0;
  color: var(--muted);
  font-size: 0.95rem;
  text-align: left;
}

.external-record-note a,
#overview a,
.source-line a,
.pdb-explorer noscript a {
  color: var(--accent-strong);
  font-weight: 700;
  text-underline-offset: 0.2em;
}

.pdb-explorer {
  margin-top: 1.25rem;
  padding: clamp(0.75rem, 2vw, 1rem);
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: var(--panel-soft);
  box-shadow: var(--shadow);
}

.pdb-toolbar {
  display: flex;
  flex-wrap: wrap;
  gap: 0.75rem;
  align-items: end;
  justify-content: space-between;
  margin-bottom: 0.875rem;
}

.field-group {
  display: grid;
  gap: 0.35rem;
  flex: 1 1 22rem;
  min-width: 0;
}

.field-group label {
  color: var(--accent-strong);
  font-size: 0.88rem;
  font-weight: 700;
}

.field-group select {
  width: 100%;
  min-height: 2.85rem;
  padding: 0.55rem 2.4rem 0.55rem 0.75rem;
  border: 1px solid var(--line);
  border-radius: var(--radius-sm);
  background: #ffffff;
  color: var(--text);
  font: inherit;
  text-overflow: ellipsis;
}

.field-group select:focus-visible,
.pdb-external-link:focus-visible {
  outline: 3px solid rgba(31, 122, 93, 0.35);
  outline-offset: 0.2rem;
}

.pdb-external-link {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 2.85rem;
  padding: 0.55rem 0.9rem;
  border-radius: var(--radius-pill);
  background: var(--accent-strong);
  color: #ffffff;
  font-weight: 700;
  text-decoration: none;
}

.pdb-layout {
  display: grid;
  grid-template-columns: minmax(0, 1.55fr) minmax(17rem, 0.45fr);
  gap: 0.875rem;
  align-items: stretch;
}

.pdb-viewer-panel,
.pdb-metadata {
  border: 1px solid var(--line);
  border-radius: var(--radius-sm);
  background: #ffffff;
  overflow: hidden;
}

.pdb-frame {
  display: block;
  width: 100%;
  min-height: clamp(34rem, 74vh, 52rem);
  border: 0;
  background: #ffffff;
}

.pdb-local-viewer {
  position: relative;
  isolation: isolate;
  overflow: hidden;
}

.pdb-local-viewer.is-static {
  display: block;
  min-height: auto;
  padding: 1rem;
  background:
    linear-gradient(180deg, rgba(247, 251, 248, 0.96), #ffffff),
    var(--panel-soft);
}

.pdb-local-viewer canvas {
  display: block;
  width: 100% !important;
  height: 100% !important;
}

.pdb-local-viewer.is-static canvas {
  display: none !important;
}

.pdb-viewer-message {
  position: absolute;
  inset: 0;
  z-index: var(--z-content, 2);
  display: grid;
  place-items: center;
  padding: 1.25rem;
  background: linear-gradient(145deg, #f7fbf8, #ffffff);
  color: var(--muted);
  font-weight: 700;
  text-align: center;
}

.pdb-viewer-message[hidden] {
  display: none;
}

.pdb-viewer-message.is-error {
  color: var(--accent-strong);
}

.pdb-viewer-message.is-static {
  position: static;
  display: grid;
  gap: 0.55rem;
  place-items: start;
  padding: 0;
  background: transparent;
  color: var(--muted);
  font-weight: 600;
  line-height: 1.55;
  text-align: left;
}

.pdb-viewer-message.is-static strong {
  color: var(--accent-strong);
  font-size: 0.78rem;
  font-weight: 800;
  letter-spacing: 0.06em;
  line-height: 1.35;
  text-transform: uppercase;
}

.pdb-viewer-message.is-static span {
  max-width: 56ch;
}

.pdb-viewer-message.is-static a {
  display: inline-flex;
  align-items: center;
  justify-content: center;
  min-height: 2.45rem;
  padding: 0.5rem 0.8rem;
  border-radius: var(--radius-pill);
  background: var(--accent-strong);
  color: #ffffff;
  font-size: 0.86rem;
  font-weight: 800;
  text-decoration: none;
}

.pdb-touch-hint {
  display: none;
  position: absolute;
  right: 0.65rem;
  bottom: 0.65rem;
  z-index: var(--z-overlay, 1);
  padding: 0.35rem 0.55rem;
  border: 1px solid rgba(255, 255, 255, 0.2);
  border-radius: var(--radius-pill);
  background: rgba(13, 79, 63, 0.86);
  color: #ffffff;
  font-size: 0.72rem;
  font-weight: 700;
  line-height: 1.2;
  pointer-events: none;
}

.pdb-metadata {
  padding: 1rem;
}

.pdb-metadata dl {
  display: grid;
  gap: 0.85rem;
}

.pdb-metadata dl div {
  padding-bottom: 0.75rem;
  border-bottom: 1px solid var(--line);
}

.pdb-metadata dt {
  color: var(--accent-strong);
  font-size: 0.78rem;
  font-weight: 700;
  letter-spacing: 0.06em;
  text-transform: uppercase;
}

.pdb-metadata dd {
  margin-top: 0.2rem;
  color: var(--muted);
  line-height: 1.5;
}

.pdb-status {
  margin: 0.85rem 0 0;
  color: var(--accent-strong);
  font-size: 0.9rem;
  font-weight: 700;
  text-align: left;
}

.guided-comparisons {
  margin-top: 1.25rem;
  padding: 1.25rem 0 0;
  border-top: 1px solid var(--line);
}

.guided-comparisons > p:not(.section-kicker) {
  max-width: 78ch;
  color: var(--muted);
}

.comparison-grid {
  display: grid;
  grid-template-columns: repeat(2, minmax(0, 1fr));
  gap: 0.875rem;
  margin-top: 1rem;
}

.comparison-card {
  padding: 1rem;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: #ffffff;
}

.comparison-index {
  margin-bottom: 0.45rem;
  color: var(--accent);
  font-size: 0.76rem;
  font-weight: 700;
  letter-spacing: 0.08em;
  text-align: left;
}

.comparison-card h4 {
  margin-bottom: 0.5rem;
  color: var(--accent-strong);
  font-size: 1.12rem;
}

.comparison-card > p:not(.comparison-index) {
  color: var(--muted);
  text-align: left;
}

.comparison-caution {
  margin: 0.75rem 0 0;
  padding-top: 0.75rem;
  border-top: 1px solid var(--line);
  font-size: 0.9rem;
  line-height: 1.5;
}

.source-line {
  margin-top: 0.85rem;
  color: var(--muted);
  font-size: 0.95rem;
  text-align: left;
}

.data-table caption {
  caption-side: top;
  margin-bottom: 0.75rem;
  text-align: left;
  color: var(--text);
  font-weight: 700;
}

.data-table th,
.data-table td {
  padding: 0.85rem;
  border: 1px solid var(--line);
  text-align: left;
  vertical-align: top;
}

.data-table thead th {
  background: var(--panel-soft);
  color: var(--accent-strong);
}

.data-table tbody th {
  width: 24%;
  background: var(--panel-soft);
  color: var(--accent-strong);
}

.workflow-list {
  display: grid;
  grid-template-columns: repeat(auto-fit, minmax(13rem, 1fr));
  gap: 0.875rem;
  margin-top: 1rem;
}

.workflow-list div {
  display: grid;
  grid-template-columns: auto 1fr;
  gap: 0.75rem;
  align-items: start;
  padding: 1rem;
  border: 1px solid var(--line);
  border-radius: var(--radius);
  background: linear-gradient(180deg, #ffffff 0%, #f7fbf8 100%);
}

.workflow-list span {
  display: inline-grid;
  place-items: center;
  width: 2rem;
  height: 2rem;
  border-radius: 50%;
  background: var(--accent);
  color: #ffffff;
  font-weight: 700;
}

.workflow-list p {
  margin: 0;
}

.learning-list {
  margin: 0;
  padding-left: 1.25rem;
}

.learning-list li {
  margin-bottom: 0.55rem;
}

.section-video {
  display: block;
  width: 100%;
  border-radius: calc(var(--radius) - 0.05rem);
  background: #08130f;
  box-shadow: inset 0 0 0 1px rgba(255, 255, 255, 0.05);
}

.figure-panel figcaption {
  margin-top: 0.75rem;
  color: var(--muted);
  line-height: 1.55;
}

.obj-preview-frame {
  display: grid;
  gap: 0;
}

.obj-preview-frame iframe {
  width: 100%;
  min-height: 34rem;
  border: 0;
  border-radius: calc(var(--radius) - 0.05rem);
  background: #040b09;
}

footer {
  padding: 1.25rem 0 2rem;
  padding-inline: clamp(1rem, 4vw, 2rem);
  border-top: 1px solid var(--line);
  color: var(--muted);
}

@media (max-width: 56rem) {
  .hero-shell,
  .section-split,
  .pdb-layout,
  .classification-layout {
    grid-template-columns: 1fr;
  }

  .page-hero {
    min-height: auto;
  }

  .hero-shell {
    padding-block: 0.85rem 1rem;
  }

  .hero-panel {
    padding: 0.85rem;
  }

  .pdb-frame {
    min-height: 32rem;
  }

  .comparison-grid {
    grid-template-columns: 1fr;
  }

  h1 {
    font-size: clamp(1.125rem, 6vw, 5rem);
  }
}

@media (max-width: 48rem) {
  main,
  footer {
    width: calc(100% - 2rem);
  }

  p,
  li,
  td,
  th,
  :where(main, .hero-panel) * {
    text-align: left;
    hyphens: none;
  }

  main p,
  main li,
  .hero-panel p,
  .figure-panel figcaption {
    width: 100%;
    max-width: none;
    margin-inline: 0;
    overflow-wrap: break-word;
    word-break: normal;
    white-space: normal;
    text-wrap: wrap;
    line-height: 1.6;
  }

  section {
    padding-block: 1.5rem;
  }

  .info-card,
  .note-panel,
  .figure-panel,
  .callout,
  .pdb-explorer,
  .comparison-card {
    min-width: 0;
    padding: 0.9rem;
  }

  .source-metadata {
    grid-template-columns: 1fr;
  }

  .classification-tree {
    overflow-x: auto;
  }

  .evolutionary-tree {
    padding: 0.85rem;
    overflow-x: hidden;
  }

  .lineage-tree,
  .lineage-tree ul {
    gap: 0.6rem;
    padding-left: 0.8rem;
  }

  .lineage-tree ul {
    margin-top: 0.6rem;
  }

  .lineage-tree li::before {
    left: -0.8rem;
    width: 0.55rem;
  }

  .lineage-node {
    grid-template-columns: minmax(0, 1fr);
    gap: 0.1rem;
    padding: 0.55rem 0.6rem;
  }

  .eyebrow,
  .section-kicker {
    font-size: 0.72rem;
    letter-spacing: 0.08em;
    line-height: 1.25;
  }

  .section-nav a {
    flex: 1 1 auto;
  }

  .pdb-toolbar,
  .pdb-external-link {
    width: 100%;
  }

  .pdb-frame {
    display: block;
    height: clamp(22rem, 62svh, 30rem);
    min-height: 22rem;
  }

  .pdb-local-viewer {
    touch-action: none;
  }

  .pdb-touch-hint {
    display: block;
  }

  .external-record {
    padding: 0.85rem;
  }

  .external-record-summary {
    grid-template-columns: 1fr;
    padding: 0.85rem;
  }

  .external-record-actions,
  .record-link {
    width: 100%;
  }

  .field-group,
  .field-group select {
    max-width: 100%;
    min-width: 0;
  }

  .field-group select {
    font-size: 0.94rem;
  }

  .data-table,
  .data-table tbody,
  .data-table thead,
  .data-table tr,
  .data-table th,
  .data-table td {
    display: block;
    width: 100%;
  }

  .data-table thead {
    display: none;
  }

  .data-table caption {
    display: block;
    width: 100%;
    padding: 0 0.15rem;
    white-space: normal;
    overflow-wrap: anywhere;
  }

  .data-table tr {
    border: 1px solid var(--line);
    border-radius: var(--radius-sm);
    margin-bottom: 0.75rem;
    overflow: hidden;
  }

  .data-table th,
  .data-table td {
    border-width: 0;
    overflow-wrap: anywhere;
    text-align: left;
    hyphens: none;
  }

  .data-table tbody th {
    border-bottom: 1px solid var(--line);
    width: 100%;
  }
}

/* Runtime fallback for mobile browsers or embedded previews with stale media-query state. */
html.is-mobile-layout main,
html.is-mobile-layout footer {
  width: calc(100% - 2rem);
}

html.is-mobile-layout main p,
html.is-mobile-layout main li,
html.is-mobile-layout .hero-panel p,
html.is-mobile-layout .figure-panel figcaption {
  width: 100%;
  max-width: none;
  margin-inline: 0;
  overflow-wrap: break-word;
  word-break: normal;
  white-space: normal;
  text-align: left;
  text-wrap: wrap;
  hyphens: none;
  line-height: 1.6;
}

html.is-mobile-layout .pdb-frame {
  display: block;
  height: clamp(22rem, 62svh, 30rem);
  min-height: 22rem;
}

html.is-mobile-layout .pdb-local-viewer {
  touch-action: none;
}

html.is-mobile-layout .pdb-touch-hint {
  display: block;
}

html.is-mobile-layout .external-record-summary {
  grid-template-columns: 1fr;
}

html.is-mobile-layout .external-record-actions,
html.is-mobile-layout .record-link {
  width: 100%;
}

html.is-mobile-layout .pdb-toolbar,
html.is-mobile-layout .pdb-external-link,
html.is-mobile-layout .field-group,
html.is-mobile-layout .field-group select {
  width: 100%;
  max-width: 100%;
  min-width: 0;
}

html.is-mobile-layout .data-table,
html.is-mobile-layout .data-table tbody,
html.is-mobile-layout .data-table thead,
html.is-mobile-layout .data-table tr,
html.is-mobile-layout .data-table th,
html.is-mobile-layout .data-table td,
html.is-mobile-layout .data-table caption {
  display: block;
  width: 100%;
}

html.is-mobile-layout .data-table thead {
  display: none;
}

html.is-mobile-layout .data-table caption,
html.is-mobile-layout .data-table th,
html.is-mobile-layout .data-table td {
  overflow-wrap: break-word;
  word-break: normal;
  text-align: left;
  hyphens: none;
}

@media (max-width: 48rem) {
  main p:not(.section-kicker):not(.comparison-index):not(.pdb-status):not(
      .source-line
    ):not(.external-record-note) {
    text-align: justify;
    text-justify: inter-word;
    hyphens: auto;
    -webkit-hyphens: auto;
    overflow-wrap: normal;
    word-break: normal;
    text-wrap: pretty;
    hyphenate-limit-chars: 6 3 3;
  }
}

html.is-mobile-layout
  main
  p:not(.section-kicker):not(.comparison-index):not(.pdb-status):not(
    .source-line
  ):not(.external-record-note) {
  text-align: justify;
  text-justify: inter-word;
  hyphens: auto;
  -webkit-hyphens: auto;
  overflow-wrap: normal;
  word-break: normal;
  text-wrap: pretty;
  hyphenate-limit-chars: 6 3 3;
}

@media (max-width: 48rem) {
  .hero-panel p,
  .hero-copy .subtitle,
  .hero-copy .hero-lead {
    width: 100%;
    max-width: none;
    text-align: justify;
    text-justify: inter-word;
    hyphens: auto;
    -webkit-hyphens: auto;
    overflow-wrap: normal;
    word-break: normal;
    white-space: normal;
    text-wrap: pretty;
    hyphenate-limit-chars: 6 3 3;
  }
}

html.is-mobile-layout .hero-panel p,
html.is-mobile-layout .hero-copy .subtitle,
html.is-mobile-layout .hero-copy .hero-lead {
  width: 100%;
  max-width: none;
  text-align: justify;
  text-justify: inter-word;
  hyphens: auto;
  -webkit-hyphens: auto;
  overflow-wrap: normal;
  word-break: normal;
  white-space: normal;
  text-wrap: pretty;
  hyphenate-limit-chars: 6 3 3;
}